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Microtubule minus ends associate with centrosomes through interactions with alpha-actinin beta-tubulin centrioles O gamma-tub

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d) gamma -tubulin

Explanation: Microtubulin emerge from MTOC(Microtubulin organising centre) which is a structure found in eukaryotic cells. Gamma tubulin is a protein which is found at centrosome. It is helpful in nucleating microtubules due to its ability to interact with tubulin monomer unit at minus end.

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