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An individual heterozygous for 4 gene pairs AaBbCcDd was testcrossed to aabbccdd, the following progeny resulted....

An individual heterozygous for 4 gene pairs AaBbCcDd was testcrossed to aabbccdd, the following progeny resulted. aBCD 42, Abcd 43, ABCd 143 abcD 142, aBcD 10, AbCd 5, ABcd 314, abCD 301. Can you please explain how to tell which gene pairs are completely linked? I am lost on this concept and have a test soon. Thank you.

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