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What does it mean to "complement (a mutation) in trans," and what is its purpose?

What does it mean to "complement (a mutation) in trans," and what is its purpose?

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Complementation does not involve genetic recombination, and therefore, the mutations must be located on two different DNA molecules (e.g., the chromosome and a plasmid), which is referred to as trans.

To perform a complementation test, two copies of the region of DNA under investigation must be present and carried on two different molecules of DNA. One copy is normally present on the chromosome and the other is carried on a second DNA molecule, typically a plasmid. For example, if we are analyzing mutants in tryptophan biosynthesis, then two copies of the whole tryptophan operon must be present. Suppose that we wish to know if two Trp- strains have a mutation in the same gene. To do this we must arrange for one mutation to be present on the chromosome and the other on a plasmid. The mutations are then referred to as being in trans with respect to one another.

Complementation test is used to describe the process to test for gene function in recessive allelism.

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